Paper · 1971 · The Journal of Agricultural Science

Plant population, shading and thinning studies in wheat

R. W. Willey; R. Holliday

11
claims
24
verbatim quotes
107
citations
1
research areas
—
Not open access

Abstract

SUMMARY Three wheat experiments are described in which a range of plant populations were shaded during different periods of development; in two of the experiments plant thinning was also carried out at a number of growth stages. Shading during the period of ear development caused an appreciable decrease in grain yield by decreasing the number of grains per ear. Shading during the grain filling period also reduced grain yield, this being brought about by decreased grain size. Thus in contrast to the barley experiments reported earlier (Willey & Holhday, 1971), these particular results gave no indication of a potential surplus of carbohydrate for grain filling and an associated limited ear capacity. However, when plant thinning was carried out at anthesis to make more carbohydrate available for grain filling in the remaining ears, grain yield per ear did not increase. It is argued, therefore, that grain yield probably was determined at least partly by a limited ear capacity. Plant thinning at earlier stages showed how the development of competition during the ear development period progressively reduced the potential capacity of the ear; the greater competition of higher plant populations accelerated this reduction in ear potential. From an examination of the effects of plant population, it is suggested that the number of grains per ear is the component having greatest influence on the decline in grain yield at above-optimum populations. The possible importance of the number of grains per unit area as an indicator of ear capacity on an area basis, and as a determinant of grain yield per unit area, is emphasized. A close relationship between grain yield per unit area and number of grains per unit area is illustrated for a number of plant-population response curves, and it is suggested that the decrease in grain yield at high populations is probably determined by a decrease in the number of grains per unit area. Evidence is presented to substantiate the idea put forward in the barley paper that this decrease in the number of grains per unit area may be attributable more to a lower production of total dry matter by the high populations during the later stages of ear development, than to an unfavourable partitioning of such dry matter between the ear and the rest of the plant.

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Claims 11

Intervention effectobservedoriginal resultnegativemeasured

Claim · model-written summary

Shading during ear development decreased wheat grain yield.

Treatment
Shading during the period of ear developmente0
Outcome
grain yielde0
Direction
decreasee0

Not reportedComparator

Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Triticum aestivum
Triticum aestivum · TaxonRasbak · CC BY-SA 3.0 · Wikimedia Commons ↗

Verbatim

e0

“Shading during the period of ear development caused an appreciable decrease in grain yield”

✓ located verbatim in the source · abstract, characters 229–319 · show in text ↑
t0

“wheat”

✓ located verbatim in the source · title, characters 50–55 · show in text ↑
Intervention effectobservedoriginal resultnegativemeasured

Claim · model-written summary

Ear development shading decreased grains per ear.

Treatment
Shading during the period of ear developmente1
Outcome
number of grains per eare0
Direction
decreasinge0

Not reportedComparator

Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Triticum aestivum
Triticum aestivum · TaxonRasbak · CC BY-SA 3.0 · Wikimedia Commons ↗

Verbatim

e0

“by decreasing the number of grains per ear.”

✓ located verbatim in the source · abstract, characters 320–363 · show in text ↑
t0

“wheat”

✓ located verbatim in the source · title, characters 50–55 · show in text ↑
e1

“Shading during the period of ear development caused an appreciable decrease in grain yield”

✓ located verbatim in the source · abstract, characters 229–319 · show in text ↑
Intervention effectobservedoriginal resultnegativemeasured

Claim · model-written summary

Grain filling shading reduced grain yield.

Treatment
Shading during the grain filling periode0
Outcome
grain yielde0
Direction
reducede0

Not reportedComparator

Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Triticum aestivum
Triticum aestivum · TaxonRasbak · CC BY-SA 3.0 · Wikimedia Commons ↗

Verbatim

e0

“Shading during the grain filling period also reduced grain yield”

✓ located verbatim in the source · abstract, characters 364–428 · show in text ↑
t0

“wheat”

✓ located verbatim in the source · title, characters 50–55 · show in text ↑
Intervention effectobservedoriginal resultnegativemeasured

Claim · model-written summary

Grain filling shading decreased grain size.

Treatment
Shading during the grain filling periode1
Outcome
grain sizee0
Direction
decreasede0

Not reportedComparator

Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Triticum aestivum
Triticum aestivum · TaxonRasbak · CC BY-SA 3.0 · Wikimedia Commons ↗

Verbatim

e0

“this being brought about by decreased grain size.”

✓ located verbatim in the source · abstract, characters 430–479 · show in text ↑
t0

“wheat”

✓ located verbatim in the source · title, characters 50–55 · show in text ↑
e1

“Shading during the grain filling period also reduced grain yield”

✓ located verbatim in the source · abstract, characters 364–428 · show in text ↑
Intervention effectobservedoriginal resultno effectmeasured

Claim · model-written summary

Thinning at anthesis did not increase yield per remaining ear.

Treatment
plant thinning was carried out at anthesise0
Outcome
grain yield per eare0
Direction
did not increasee0

Not reportedComparator

Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Triticum aestivum
Triticum aestivum · TaxonRasbak · CC BY-SA 3.0 · Wikimedia Commons ↗

Verbatim

e0

“when plant thinning was carried out at anthesis to make more carbohydrate available for grain filling in the remaining ears, grain yield per ear did not increase.”

✓ located verbatim in the source · abstract, characters 716–878 · show in text ↑
t0

“wheat”

✓ located verbatim in the source · title, characters 50–55 · show in text ↑
Observationobservedoriginal resultnegativemeasured

Claim · model-written summary

Earlier thinning showed competition progressively reduced ear capacity.

Outcome
potential capacity of the eare0
Direction
reducede0
Qualifier
competition during the ear development periode0

Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Triticum aestivum
Triticum aestivum · TaxonRasbak · CC BY-SA 3.0 · Wikimedia Commons ↗

Verbatim

e0

“Plant thinning at earlier stages showed how the development of competition during the ear development period progressively reduced the potential capacity of the ear”

✓ located verbatim in the source · abstract, characters 988–1152 · show in text ↑
t0

“wheat”

✓ located verbatim in the source · title, characters 50–55 · show in text ↑
Associationobservedoriginal resultnegativemeasured

Claim · model-written summary

Higher population competition accelerated reduction in ear potential.

Outcome
ear potentiale0
Direction
reductione0
Setting
higher plant populationse0

Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Triticum aestivum
Triticum aestivum · TaxonRasbak · CC BY-SA 3.0 · Wikimedia Commons ↗

Verbatim

e0

“the greater competition of higher plant populations accelerated this reduction in ear potential.”

✓ located verbatim in the source · abstract, characters 1154–1250 · show in text ↑
t0

“wheat”

✓ located verbatim in the source · title, characters 50–55 · show in text ↑
Associationauthor interpretationoriginal resultmeasured

Claim · model-written summary

Grains per ear were suggested to most influence yield decline above optimum population.

Organ / stage
eare0
Outcome
grain yielde0
Direction
declinee0
Qualifier
above-optimum populationse0

Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Triticum aestivum
Triticum aestivum · TaxonRasbak · CC BY-SA 3.0 · Wikimedia Commons ↗

Verbatim

e0

“the number of grains per ear is the component having greatest influence on the decline in grain yield at above-optimum populations.”

✓ located verbatim in the source · abstract, characters 1328–1459 · show in text ↑
t0

“wheat”

✓ located verbatim in the source · title, characters 50–55 · show in text ↑
Associationobservedoriginal resultmeasured

Claim · model-written summary

Grain yield per area closely related to grains per area across population response curves.

Outcome
grain yield per unit areae0
Setting
plant-population response curvese0

Not reportedDirection

Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Triticum aestivum
Triticum aestivum · TaxonRasbak · CC BY-SA 3.0 · Wikimedia Commons ↗

Verbatim

e0

“A close relationship between grain yield per unit area and number of grains per unit area is illustrated for a number of plant-population response curves”

✓ located verbatim in the source · abstract, characters 1638–1791 · show in text ↑
t0

“wheat”

✓ located verbatim in the source · title, characters 50–55 · show in text ↑
Mechanismauthor interpretationoriginal resultmeasured

Claim · model-written summary

High population yield decline was suggested to result from fewer grains per area.

Outcome
decrease in grain yielde0
Qualifier
decrease in the number of grains per unit areae0

Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Triticum aestivum
Triticum aestivum · TaxonRasbak · CC BY-SA 3.0 · Wikimedia Commons ↗

Verbatim

e0

“the decrease in grain yield at high populations is probably determined by a decrease in the number of grains per unit area.”

✓ located verbatim in the source · abstract, characters 1818–1941 · show in text ↑
t0

“wheat”

✓ located verbatim in the source · title, characters 50–55 · show in text ↑
Mechanismauthor interpretationoriginal resultmeasured

Claim · model-written summary

Fewer grains per area at high density were attributed more to reduced late dry matter production than unfavorable partitioning.

Outcome
decrease in the number of grains per unit areae0
Qualifier
lower production of total dry mattere0

Organisms pictured · Representative pictures of the matched species, not figures from the paper.

Triticum aestivum
Triticum aestivum · TaxonRasbak · CC BY-SA 3.0 · Wikimedia Commons ↗

Verbatim

e0

“this decrease in the number of grains per unit area may be attributable more to a lower production of total dry matter by the high populations during the later stages of ear development, than to an unfavourable partitioning of such dry matter between the ear and the rest of the plant.”

✓ located verbatim in the source · abstract, characters 2026–2311 · show in text ↑
t0

“wheat”

✓ located verbatim in the source · title, characters 50–55 · show in text ↑

shade · yield · Triticum aestivum (wheat)Positive and negative results

69 papers: 9 positive, 60 negative or no effect

59 papers in the same direction as this paper · 9 in the opposite direction

  1. negative
  2. negative
  3. no effect

65 papers more in this group

See all 69 papers →

pruning · yield · Triticum aestivum (wheat)Positive and negative results

93 papers: 33 positive, 60 negative or no effect

59 papers in the same direction as this paper · 33 in the opposite direction

  1. positive
  2. positive
  3. positive

89 papers more in this group

See all 93 papers →

Claims are grouped by organism, treatment and what was measured or targeted. A paper counts as positive when any of its claims in the group is positive.